Suidae

Suidae; Temporal range: Oligocene–Holocene PreꞒ Ꞓ O S D C P T J K Pg N
	Suid species of different genera; from top-left, clockwise: Wild boar (Sus scrofa), pygmy hog (Porcula salvania), giant forest hog (Hylochoerus meinertzhageni), red river hog (Potamochoerus porcus), common warthog (Phacochoerus africanus), North Sulawesi babirusa (Babyrousa celebensis)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Artiodactyla
Suborder:	Suina
Family:	Suidae; Gray, 1821
Type genus
	Sus; Linnaeus, 1758
Genera
	Over 30 extinct genera, 6 extant, ; see text.

Suidae is a family of artiodactyl mammals which are commonly called pigs, hogs, or swine. In addition to numerous fossil species, 18 extant species are currently recognized (or 19 counting domestic pigs and wild boars separately), classified into between four and eight genera. Within this family, the genus Sus includes the domestic pig, Sus scrofa domesticus or Sus domesticus, and many species of wild pig from Europe to the Pacific. Other genera include babirusas and warthogs. All suids, or swine, are native to the Old World, ranging from Asia to Europe and Africa.

The earliest fossil suids date from the Oligocene epoch in Asia, and their descendants reached Europe during the Miocene.^[1] Several fossil species are known and show adaptations to a wide range of different diets, from strict herbivory to possible carrion-eating (in Tetraconodontinae).^[2]

Physical characteristics

Suids belong to the order Artiodactyla, and are generally regarded as the living members of that order most similar to the ancestral form. Unlike most other members of the order, they have four toes on each foot, although they walk only on the middle two digits, with the others staying clear of the ground. They also have a simple stomach, rather than the more complex ruminant stomach found in most other artiodactyl families.^[3]

They are small to medium animals, varying in size from 58 to 66 cm (23 to 26 in) in length, and 6 to 9 kg (13 to 20 lb) in weight in the case of the pygmy hog, to 130–210 cm (4.3–6.9 ft) and 100–275 kg (220–606 lb) in the giant forest hog.^[4] They have large heads and short necks, with relatively small eyes and prominent ears. Their heads have a distinctive snout, ending in a disc-shaped nose. Suids typically have a bristly coat, and a short tail ending in a tassle.^{[citation needed]} The males possess a corkscrew-shaped penis, which fits into a similarly shaped groove in the female's cervix.^[5]^[6]^[7]

Suids have a well-developed sense of hearing, and are vocal animals, communicating with a series of grunts, squeals, and similar sounds. They also have an acute sense of smell. Many species are omnivorous, eating grass, leaves, roots, insects, worms, and even frogs or mice. Other species are more selective and purely herbivorous.^[3]

Their teeth reflect their diet, and suids retain the upper incisors, which are lost in most other artiodactyls. The canine teeth are enlarged to form prominent tusks, used for rooting in moist earth or undergrowth, and in fighting. They have only a short diastema. The number of teeth varies between species, but the general dental formula is: 1–3.1.2–4.3030.1.020.3.

Behavior and reproduction

Suids are intelligent and adaptable animals. Adult females (sows) and their young travel in a group (sounder; see List of animal names), while adult males (boars) are either solitary, or travel in small bachelor groups. Males generally are not territorial, and come into conflict only during the mating season.

Litter size varies between one and twelve, depending on the species. The mother prepares a grass nest or similar den, which the young leave after about ten days. Suids are weaned at around three months, and become sexually mature at 18 months. In practice, however, male suids are unlikely to gain access to sows in the wild until they have reached their full physical size, at around four years of age. In all species, the male is significantly larger than the female, and possesses more prominent tusks.^[3]

Classification

The following 18 extant species of suid are currently recognised:^[8]

Image	Genus	Living species
	Sus – pigs	Sus ahoenobarbus – Palawan bearded pig Sus barbatus – Bornean bearded pig Sus cebifrons – Visayan warty pig Sus celebensis – Sulawesi warty pig Sus oliveri – Mindoro warty pig Sus philippensis – Philippine warty pig Sus scrofa – wild boar Sus scrofa domesticus – domestic pig (sometimes treated as a full species, S. domestica) Sus verrucosus – Javan warty pig
	Porcula	Porcula salvania – pygmy hog
	Hylochoerus	Hylochoerus meinertzhageni – giant forest hog
	Potamochoerus	Potamochoerus larvatus – bushpig Potamochoerus porcus – red river hog
	Phacochoerus – warthog	Phacochoerus africanus – common warthog Phacochoerus aethiopicus – desert warthog
	Babyrousa – babirusa	Babyrousa babyrussa – Moluccan babirusa Babyrousa bolabatuensis – Bola Batu babirusa Babyrousa celebensis – North Sulawesi babirusa Babyrousa togeanensis – Togian babirusa

Phylogeny

Cladogram of Suidae. Mikko's Phylogeny Archive (Based is McKenna & Bell, 1997, Liu, 2003 и Harris & Liu, 2007):^[9]

? † Mabokopithecus

† Hemichoerus

† Paradoxodonides

† Cainochoerus

† Hyotheriinae

	† Chleuastochoerus

	† Dubiotherium

	† Sinapriculus

	† Hyotherium

† Listriodontinae

	† Listriodon

	† Lopholistriodon

	† Eurolistriodon

	† Bunolistriodon

† Kubanochoerinae

	† Nguruwe

	? † Kenyasus

	† Libycochoerus

	† Megalochoerus

	† Kubanochoerus

† Miochoerus

† Tetraconodontinae

† Conohyus

† Tetraconodon

† Parachleuastochoerus

† Lophochoerus

† Sivachoerus

	† Nyanzachoerus

	† Notochoerus

† Namachoerinae

† Namachoerus

Suinae

Suini

† Korynochoerus

† Hippopotamodon

† Eumaiochoerus

† Microstonyx

Sus sensu lato

	Sus sensu stricto

	Porcula

	? Dasychoerus

	? Euhys

Potamochoerini

? † Celebochoerus

? † Propotamochoerus

† "Kolpochoerus" afarensis

Potamochoerus

	† Kolpochoerus

	Hylochoerus

† Hippohyini

	† Sivahyus

	† Hippohyus

	? † Sinohyus

Babyrousini

Babyrousa

Phacochoerini

	† Potamochoeroides

	† Metridiochoerus

	Phacochoerus

References

^ Palmer, D., ed. (1999). The Marshall Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals. London: Marshall Editions. p. 269. ISBN 1-84028-152-9.
^ Savage, R. J. G.; Long, M. R. (1986). Mammal Evolution: an illustrated guide. New York: Facts on File. pp. 212–213. ISBN 0-8160-1194-X.
^ ^a ^b ^c Cumming, David (1984). Macdonald, D. (ed.). The Encyclopedia of Mammals. New York: Facts on File. pp. 500–503. ISBN 0-87196-871-1.
^ "Forest hog (Hylochoerus meinertzhageni) - Quick facts".
^ Bonnie S. Dunbar; M.G. O'Rand (29 June 2013). A Comparative Overview of Mammalian Fertilization. Springer Science & Business Media. pp. 330–. ISBN 978-1-4757-8982-9.
^ Peter G. G. Jackson; Peter D. Cockcroft (2007). Handbook of Pig Medicine. Elsevier Health Sciences. ISBN 978-0-7020-2828-1.
^ Virginia Douglass Hayssen; Ari Van Tienhoven (1993). Asdell's Patterns of Mammalian Reproduction: A Compendium of Species-specific Data. Cornell University Press. ISBN 0-8014-1753-8. suidae penis.
^ Wilson, Don E.; Mittermeier, Russell A., eds. (2011). Handbook of the Mammal Species of the World, vol. 2. Barcelona: Lynx Edicions. pp. 274–291. ISBN 978-8496553774.
^ "SUIDAE – pigs". Mikko's Phylogeny Archive. Archived from the original on 2021-07-17. Retrieved 2021-08-13.

External links

Data related to Suidae at Wikispecies Media related to Suidae at Wikimedia Commons

[1] Palmer, D., ed. (1999). The Marshall Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals. London: Marshall Editions. p. 269. ISBN 1-84028-152-9.

[MamEv-2] Savage, R. J. G.; Long, M. R. (1986). Mammal Evolution: an illustrated guide. New York: Facts on File. pp. 212–213. ISBN 0-8160-1194-X.

[EoM-3] Cumming, David (1984). Macdonald, D. (ed.). The Encyclopedia of Mammals. New York: Facts on File. pp. 500–503. ISBN 0-87196-871-1.

[4] "Forest hog (Hylochoerus meinertzhageni) - Quick facts".

[DunbarO'Rand2013-5] Bonnie S. Dunbar; M.G. O'Rand (29 June 2013). A Comparative Overview of Mammalian Fertilization. Springer Science & Business Media. pp. 330–. ISBN 978-1-4757-8982-9.

[JacksonCockcroft2007-6] Peter G. G. Jackson; Peter D. Cockcroft (2007). Handbook of Pig Medicine. Elsevier Health Sciences. ISBN 978-0-7020-2828-1.

[HayssenTienhoven1993-7] Virginia Douglass Hayssen; Ari Van Tienhoven (1993). Asdell's Patterns of Mammalian Reproduction: A Compendium of Species-specific Data. Cornell University Press. ISBN 0-8014-1753-8. suidae penis.

[8] Wilson, Don E.; Mittermeier, Russell A., eds. (2011). Handbook of the Mammal Species of the World, vol. 2. Barcelona: Lynx Edicions. pp. 274–291. ISBN 978-8496553774.

[9] "SUIDAE – pigs". Mikko's Phylogeny Archive. Archived from the original on 2021-07-17. Retrieved 2021-08-13.

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